Players have the ability to roam anywhere on the map, or leave a conversation at any time. They have the ability to customize weapons; the game includes over 50 base guns, which can be crafted with a variety of modifications, such as barrel types and laser focus, with over 700 modifications available. Power Armor has been redesigned to be more like a vehicle than an equipable suit of armor, and can also be modified, adding items such as a jetpack or selecting separate types of armor for each part of the suit. A new feature to the series is the ability to craft and deconstruct settlements and buildings. Players can select some in-game objects and structures, and use them to freely build their own structures. Players can build various defenses around their settlements, such as turrets and traps, to defend against random attacks.

At the beginning of the game, players are given points to spend on a character progression system called S.P.E.C.I.A.L.. The system represents seven statistics, namely strength, perception, endurance, charisma, intelligence, agility and luck. When players gain enough experience points to gain a new level, they unlock a new ability. When players allocate more points to a stat, more abilities can be unlocked. These perks can also be upgraded to improve the protagonists efficiency and to further unlock new abilities. There are about 275 perks available for players to unlock. There is no level cap and the game does not end once the main story is complete.
The series also allows for the player to have a companion character follow and assist them, similar in style to Fallout 3 and Fallout: New Vegas. There are thirteen possible companions, seven of which (Dogmeat, Codsworth, Preston Garvey, Piper Wright, Nick Valentine, John Hancock, and Deacon) must be encountered during the main quests, although only Dogmeat is required to join. For the first time in the series, these followers can interact with the environment on the player characters behalf. For example, if the player character does not have required skills to hack a terminal or pick a lock, they can order the companion to do it for them. The player may only travel with one companion at a time, although the player is accompanied by other characters in certain quests. Some of the companions can be romanced by the player.
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